Ministry of Education Key Laboratory for Ecology of Tropical Islands College of Life Sciences Hainan Normal University Haikou 571158 China;
Ministry of Education Key Laboratory for Ecology of Tropical Islands College of Life Sciences Hainan Normal University Haikou 571158 China;
Ministry of Education Key Laboratory for Ecology of Tropical Islands College of Life Sciences Hainan Normal University Haikou 571158 China;
Laboratoired'Ecologie Systématique et Evolution CNRS UMR 8079 Université Paris-Sud Batiment 362 Orsay Cedex F-91405 France;
Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering College of Life Sciences Beijing Normal University Beijing 100875 China;