Key Laboratory of Aquatic Botany and Watershed Ecology;
Wuhan Botanical Garden;
Chinese Academy of Sciences;
Wuhan 430074;
P.R.China;
College of Bioscience and Biotechnology;
Hunan Agricultural University;
Changsha 410128;
P.R. China;
Graduate University;
Chinese Academy of Sciences;
Beijing 100049;
P.R. China;