School of Life Sciences Guizhou University/State Engineering Technology Institute for Karst Desertification Control/Key Laboratory of Plant Resources Conservation and Germplasm Innovation in Mountainous Region Ministry of Education Guiyang 550025 P.R.China;
Special Key Laboratory of Microbial Resources and Drug Development from Higher Education Institution of Guizhou Province/ Research Center for Medicine and Biology Zunyi Medical University Zunyi 563006 P.R.China;
School of Agriculture Guizhou University/Guizhou Sub-Center of National Wheat Improvement Center Guiyang 550025 P.R.China;