College of Plant Protection Shandong Agricultural University Tai'an 271018 PR. China;
State Key Laboratory for Biology of Plant Disease and Insect Pests Institute of Plant Protection Chinese Academy of Agricultural Sciences Beijing 100193 P.R. China;
Functional and Evolutionary Entomology Gembloux Agro-Bio Tech University of Liege Passage des DOportds 2 Gembloux B-5030 Belgium;