Guangdong Provincial Key Laboratory of Fishery Ecology and Environment;
Key Laboratory of South China Sea Fishery Resources Development and Utilization of Ministry of Agriculture;
South China Sea Fisheries Research Institute;
Chinese Academy of Fishery Sciences;
Guangzhou 510300;
P. R. China;
Key Laboratory of Tropical Marine Bio-resources and Ecology;
South China Sea Institute of Oceanology;
Chinese Academy of Sciences;
Guangzhou 510301;
P. R. China;