[1]Hunan Academy of Forestry Hengshan Research Station of Forest Ecosystem Changsha 410004 China;
[2]School of Science Central South University of Forestry and Technology Changsha 410004 China;
[3]Hunan Hengshan National Nature Reserve Administration Nanyue 421999 China;
[4]School of Life and Health Science Kaili University Kaili 556011 China;
[5]Water Resources Research & Development Center Changsha 410013 China;
Evergreen broad-leaved forest; Carbon reserve of arbor layer; Mid-subtropical zone;