Key Laboratory of Economic Plants and Biotechnology Yunnan Key Laboratory for Wild Plant Resources Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 People's Republic of China;
University of Chinese Academy of Sciences Beijing 100049 People's Republic of China;
Key Laboratory of Economic Plants and Biotechnology Yunnan Key Laboratory for Wild Plant Resources Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 People's Republic of China;