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New Eocene Coleoid (Cephalopoda) Diversity from Statolith Remains: Taxonomic Assignation Fossil Record Analysis and New Data for Calibrating Molecular Phylogenies

机译:从石蜡中获得的新始新世胶质(Cephalopoda)多样性:分类分配化石记录分析和用于校准分子系统发育的新数据

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摘要

New coleoid cephalopods are described from statolith remains from the Middle Eocene (Middle Lutetian) of the Paris Basin. Fifteen fossil statoliths are identified and assigned to the Sepiidae (Sepia boletzkyi sp. nov.,? Sepia pira sp. nov.), Loliginidae (Loligo clarkei sp. nov.), and Ommastrephidae (genus indet.) families. The sediments containing these fossils indicate permanent aquatic settings in the infralittoral domain. These sediments range in age from 46 Mya to 43 Mya. Analysis of the fossil record of statoliths (from findings described here, together with a review of previously published data) indicates marked biases in our knowledge. Fossil statoliths are known from as far back as the Early Jurassic (199.3 to 190.8 Mya) but surprisingly, to the best of our knowledge, no record occurs in the Cretaceous. This is a “knowledge bias” and clearly calls for further studies. Finally, we attempt to compare findings described here with fossils previously used to constrain divergence and/or diversification ages of some coleoid subclades in molecular phylogenies. This comparison clearly indicates that the new records detailed here will challenge some estimated divergence times of coleoid cephalopod subclades.
机译:从巴黎盆地中始新世(Lutetian中部)的层状石遗骸中描述了新的类脂头足类。鉴定出15个化石层状石并分配给Sepiidae(Sepia boletzkyi sp。nov。,?Sepia pira sp。nov。),Loliginidae(Loligo clarkei sp。nov。)和Ommastrephidae(属indet。)科。含有这些化石的沉积物表明在下层域有永久的水生环境。这些沉积物的年龄范围从46 Mya到43 Mya。对层状石化石记录的分析(来自此处描述的发现,以及对先前发布的数据的回顾)表明,我们的知识存在明显偏差。化石针石是最早的侏罗纪(199.3至190.8 Mya),但令人惊讶的是,据我们所知,白垩纪没有记录。这是“知识偏见”,显然需要进一步研究。最后,我们尝试将此处描述的发现与以前用于限制分子系统发育中一些类胚体子瓣的发散和/或发散年龄的化石进行比较。这种比较清楚地表明,此处详细介绍的新记录将挑战类脂头足类亚群的某些估计发散时间。

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