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The Tangled Nature model with inheritance and constraint: Evolutionary ecology restricted by a conserved resource

机译：具有继承性和约束性的纠结自然模型：受保守资源约束的进化生态学

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摘要

We study a version of the Tangled Nature model of evolutionary ecology redefined in a phenotype space where mutants have properties correlated to their parents. The model has individual-based dynamics whilst incorporating species scale competitive constraints and a system scale resource constraint. Multiple species arise that coexist in a species interaction network with evolving global properties. Both the mean interaction strength and the network connectance increase relative to the null system as mutualism becomes more extensive. From a study of the dependence of average degree on the resource level we extract the diversity-connectance relationship which conforms to the hyperbolic form seen in field data. This is adjudged to arise as a consequence of the evolutionary pressure to achieve positive interactions. The network degree distributions conform more strongly to exponential than to the null binomial distributions in all cases. This effect is believed to be caused by correlations in the reproductive process. We also study how resource availability influences the phenotypical lifetime distribution which is approximately of power law form. We observe that the mean lifetime is inversely related to the resource level.
机译：我们研究了在表型空间中重新定义的进化生态的纠结自然模型的一个版本，在该表型空间中，突变体具有与其父母相关的特性。该模型具有基于个体的动力学，同时结合了物种规模的竞争约束和系统规模的资源约束。多种物种共存于具有不断发展的全球特性的物种相互作用网络中。随着互惠关系的发展，相对于零系统，平均交互强度和网络连接性均增加。通过研究平均程度对资源水平的依赖性，我们提取了多样性-联系关系，该关系与现场数据中的双曲线形式相符。可以断定，这是实现积极互动的进化压力的结果。在所有情况下，网络度分布与指数的相合性要比零二项式分布更强。据信这种作用是由生殖过程中的相关性引起的。我们还研究了资源可用性如何影响表型寿命分布，该寿命分布近似于幂律形式。我们观察到平均寿命与资源水平成反比。

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