首页> 外文期刊>Journal of Systematic Palaeontology >First cranial remains of Cheirogaster richardi (Testudines: Testudinidae) from the Late Miocene of Ecoparc de Can Mata (Valles-Penedes Basin, NE Iberian Peninsula): taxonomic and phylogenetic implications
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First cranial remains of Cheirogaster richardi (Testudines: Testudinidae) from the Late Miocene of Ecoparc de Can Mata (Valles-Penedes Basin, NE Iberian Peninsula): taxonomic and phylogenetic implications

机译:Ecoparc de Can Mata(中伊比利亚半岛瓦莱-佩内德斯盆地)的中新世晚期的Cheirogaster richardi(睾丸科:Testudinidae)的头颅遗骸:分类学和系统发育意义

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摘要

Although skulls of extinct Testudinidae are generally much scarcer than shell remains, when available they provide important data for resolving taxonomic and phylogenetic problems, as illustrated here by two well-preserved giant tortoise skulls from the early Vallesian (MN9, Late Miocene) of Ecoparc de Can Mata (ECM; els Hostalets de Pierola, Valles-Penedes Basin, NE Iberian Peninsula). These specimens, referable to the extinct genus Cheirogaster, differ significantly from C. bolivari and are assigned to C. richardi, whose cranial morphology was previously unknown. This nominal taxon had been considered a junior subjective synonym of C. bolivari, due to a previous neotype designation for the former, based on shell remains, that did not meet the requirements of the International Code of Zoological Nomenclature. This designation is here considered invalid because it was based on material from a different geographical area, even though remains from the original type locality area were available. Given that the holotype of C. richardi (from the early Vallesian of els Hostalets de Pierola) has been destroyed, to clarify the taxonomic status of this taxon we here designate one of the two ECM skulls as the neotype of the species. On this basis, an emended diagnosis is provided, which leads us to conclude that two different species are recorded from the Iberian Miocene: C. bolivari, from the middle Aragonian of the inner Iberian basins; and C. richardi, from the latest Aragonian and Vallesian of the Valles-Penedes Basin. Additional cranial material of Cheirogaster from inner Iberia would be required to clarify whether these species display distinct geographical distributions and/or different chronostratigraphical ranges. A cladistic analysis of Testudinidae based on cranial morphology supports a sister-taxon relationship between Cheirogaster and Centrochelys. Overall, our results highlight the significance of cranial morphology for attaining a better understanding of turtle taxonomy and phylogenetic relationships.
机译:尽管已灭绝的Testudinidae颅骨通常比贝壳残骸稀少得多,但在可用时,它们提供了解决分类学和系统发育问题的重要数据,如此处所示,这是两个保存良好的Ecoparc de早期瓦利西亚(MN9,中新世晚期)乌龟头骨的例证。 Can Mata(ECM;伊​​比利亚半岛东北部Valles-Penedes盆地的els Hostalets de Pierola)。这些标本与已灭绝的白尾蛇属(Cheirogaster)有关,与玻利虫(C. bolivari)有显着差异,并被分配给C. richardi,后者的颅骨形态以前未知。由于以前的基于壳残留物的新类型名称不符合《国际动物学命名法》的要求,因此该名义生物分类被认为是玻里氏梭菌的初级主观同义词。此名称在这里被认为是无效的,因为它是基于来自不同地理区域的资料,即使可以得到原始类型区域的残留物。鉴于C. richardi的原型(来自els Hostalets de Pierola的早期瓦利西亚人)已被摧毁,为了阐明该类群的分类学地位,在此我们将两个ECM头骨之一指定为该物种的新类型。在此基础上,提供了一种改进的诊断方法,这使我们得出结论:伊比利亚中新世记录了两个不同的物种:C. bolivari,来自伊比利亚内部盆地的中部阿拉贡。和C. richardi,来自Valles-Penedes盆地的最新Aragonian和Vallesian。需要从伊比利亚内陆获得更多的白尾蛇颅骨材料,以阐明这些物种是否显示出独特的地理分布和/或不同的年代地层范围。基于颅骨形态的睾丸科的分类分析支持了Cheirogaster和Centrochelys之间的姐妹-分类单元关系。总体而言,我们的结果突出了颅骨形态对于更好地了解乌龟分类学和系统发育关系的重要性。

著录项

  • 来源
    《Journal of Systematic Palaeontology》 |2014年第8期|833-864|共32页
  • 作者单位

    Institut Catala de Paleontologia, Universitat Autonoma de Barcelona. Edifici ICP, Campus de la UAB s, 08193 Cerdanyola del Valles, Barcelona, Spain;

    Institut Catala de Paleontologia, Universitat Autonoma de Barcelona. Edifici ICP, Campus de la UAB s, 08193 Cerdanyola del Valles, Barcelona, Spain,Dipartimento di Scienze della Terra, Universita degli Studi di Torino, Via Valperga Caluso 35,I-10125 Torino, Italy;

    Institut Catala de Paleontologia, Universitat Autonoma de Barcelona. Edifici ICP, Campus de la UAB s, 08193 Cerdanyola del Valles, Barcelona, Spain;

    Institut Catala de Paleontologia, Universitat Autonoma de Barcelona. Edifici ICP, Campus de la UAB s, 08193 Cerdanyola del Valles, Barcelona, Spain,FOSSILIA Serveis Paleontologics i Geologics, S.L. c/Jaume I num. 87, Ier 5a, 08470 Sant Celoni, Barcelona, Spain;

    Dipartimento di Scienze della Terra, Universita degli Studi di Torino, Via Valperga Caluso 35,I-10125 Torino, Italy,Institut Catala de Paleontologia, Universitat Autonoma de Barcelona. Edifici ICP, Campus de la UAB s, 08193 Cerdanyola del Valles, Barcelona, Spain;

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  • 正文语种 eng
  • 中图分类
  • 关键词

    giant tortoises; testudinids; taxonomy; phylogeny; Cheirogaster bolivari; early Vallesian;

    机译:乌龟;睾丸;分类;系统发育Cheirogaster bolivari;早期的瓦利西亚人;

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