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首页> 外文期刊>Journal of Systematic Palaeontology >Phylogenetic reappraisal of Rhytidosteidae (Stereospondyli: Trematosauria), temnospondyl amphibians from the Permian and Triassic
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Phylogenetic reappraisal of Rhytidosteidae (Stereospondyli: Trematosauria), temnospondyl amphibians from the Permian and Triassic

机译：对二叠纪和三叠纪的Rhytidosteidae（Stereospondyli：Trematosauria），temnospondyl两栖动物进行系统发育的重新评估。

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All stereospondyl taxa previously placed within Rhytidosteidae are reviewed in a phylogenetic context. A parsimony analysis shows that Pneumatostega, Trucheosaurus, Rhytidosieus and Nanolania are placed outside Rhytidosteidae. If an implied weighting method is introduced in order to downweight homoplastic characters, then these taxa are placed within rhytidosteids (with the exception of Rhytidosteus), thus forming a resolved monophyletic group. This group is supported by four synapomorphies, including a twisted pterygoid quadrate ramus and sculpture consisting of a reticulate pattern with pustules or nodules at the junction between adjacent crests and ridges. Based on the topology of a majority rule consensus tree, we postulate a close relationship among all eastern Gondwanan rhytidosteids (Australian and Indian taxa) and among western Gondwanan taxa (South America, South Africa and Madagascar). Laurasian Boreopelta and Peltostega are successive paraphyletic taxa of both eastern and western groups. The Australian family 'Derwentiidae' is nested within Rhytidosteidae and is redefined as a subfamily (Derwentiinae nov.) which also includes Indobrachyops. Peltosteginae and Indobrachyopidae as currently defined in the literature are not supported. Rhytidosteinae is also not supported because Laidleria, Pneumatostega and Rhytidosteus do not emerge as a clade. The presence of the rhytidosteid Trucheosaurus major in the Late Permian of Australia suggests that rhytidosteids were already diversified at that time. Increasing efforts to collect Upper Permian and Lower Triassic rhytidosteids should help to improve stratigraphical, palaeogeographical and phylogenetic information regarding this widespread group of temnospondyls.
机译：以前放置在Rhydidosteidae中的所有立体骨分类群都在系统发育背景下进行了综述。简化分析表明，肺炎科，真龙属，假单胞菌和纳兰氏菌被放置在Rhytidosteidae外。如果引入隐式加权方法以降低同质性，则将这些类群放在Rhytidosteids（Rhytidosteus除外）内，从而形成一个解析的单系群。该族群由四个同形亚种所支撑，包括扭曲的翼龙四角形支和由网状图案组成的雕塑，在相邻波峰与山脊之间的交界处有脓疱或结节。基于多数规则共识树的拓扑结构，我们假设所有东部的冈瓦南类Rhytidosteids（澳大利亚和印度的分类单元）与西部的冈瓦纳类群（南美，南非和马达加斯加）之间存在密切的关系。 Laurasian Boreopelta和Peltostega是东方和西方群体相继的寄生虫类群。澳大利亚的“ Derwentiidae”科嵌套在Rhytidosteidae内，被重新定义为亚科（Derwentiinae nov。），其中还包括Indobrachyops。不支持目前文献中所定义的猪角鲨科和支气管龙科。不支持Rhytidosteinae，因为Laidleria，Pneumatostega和Rhytidosteus不作为进化枝出现。在澳大利亚的二叠纪晚期，存在着节肢动物Trucheosaurus专业，这表明当时的节肢动物已经多样化。加大收集上二叠纪和下三叠纪的类胡萝卜素的努力，应有助于改善有关这一广泛的类脊柱石类的地层，古地理和系统发育信息。

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