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The origin and early evolution of the osteostraci (vertebrata): a phylogeny for the thyestiida

机译:骨纹层(椎骨)的起源和早期演化:胸腺科的系统发育

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摘要

The Osteostraci are currently regarded as the sister taxon to jawed vertebrates and, as such, osteostracan phytogeny, notably their hypothetical ancestral morphotype, has far reaching ramifications for early vertebrate evolution. The polarity of the group and the monophyly and interrelationships of the proposed constituent clades have not been tested. Here a new phylogeny is proposed for the Thyestiida, the most controversial and informative of the five main orders of Osteostraci. A comprehensive global parsimony approach is applied for the first time. The monophyly of the group, including genera of suspected thyestiid affinities, is firmly demonstrated, as is the monophyly of some constituent groups such as the Siberian Tannuaspidinae. The phylogeny supports the proposals of Janvier (1985a-c) and is inconsistent with previous 'Tremataspis-basal' models of Osteostracan evolution. Application of the phylogeny to the stratigraphical record demonstrates an early evolution and origin for the Osteostraci and indicates a missing fossil record for the group in the Llandovery, strongly related to facies bias.
机译:Osteostraci目前被认为是有颚脊椎动物的姊妹分类群,因此,stratrastracan的植物遗传学,特别是其假想的祖先形态,对脊椎动物的早期进化产生了深远的影响。该组的极性以及拟议组成进化枝的单亲和相互关系尚未经过测试。在这里,提出了一种新的系统发育学,用于胸腺科,这是骨成骨纲五个主要科目中最具争议和最有益的内容。首次使用全面的全局简约方法。包括可疑的类甲状腺亲和性属在内的这一类的单亲生物得到了可靠的证明,某些组成群如西伯利亚Tannuaspidinae的单亲生物也得到了证实。系统发育支持Janvier(1985a-c)的建议,并且与以前的骨成骨动物进化的'Tremataspis-basal'基础模型不一致。系统发育学在地层记录中的应用证明了骨层古迹的早期演化和起源,并表明在兰德奥弗里该组的化石记录缺失,这与相偏差密切相关。

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