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The oldest known cyclophoroidean land snails (Caenogastropoda) from Asia

机译:来自亚洲的最古老的环纹陆蜗牛(Caenogastropoda)

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摘要

The earliest Miocene (Aquitanian, 23-21 Ma) Hang Mon Formation at Hang Mon in Northern Vietnam has yielded a rich assemblage of terrestrial gastropods. Four species from this assemblage belong to the land caenogastropod superfamily Cyclophoroidea. Three of these are assigned to genera with Recent representatives in Southeast Asia and are described as new species: Cyclophorus hangmonensis Raheem & Schneider sp. nov. (Cyclophoridae: Cyclophorini), Alycaeus sonlaensis Raheem & Schneider sp. nov. (Cyclophoridae: Alycaeinae) and Tortulosa naggsi Raheem & Schneider sp. nov (Pupinidae: Pupinellinae). These fossil species represent the earliest records for their genera and are thus of great value for calibrating molecular phylogenies of the Cyclophoroidea. The fourth species is represented only by poorly preserved fragments and is retained in open nomenclature in the Cyclophoridae. While extant Cyclophoroidea have their greatest diversity in Tropical Asia, the oldest fossils described to date from the region are from the Late Pleistocene. The fossils from Hang Mon predate these by more than 20 million years and are also globally among the earliest cyclophoroideans recorded from the area of the present-day forested Tropics. Cyclophoroidea older than the Miocene are only known from Europe. Given that Recent Cyclophoridae and Pupinidae are typically associated with tropical forests, it seems likely that the Hang Mon fossils were also tropical forest taxa and that suitable habitat was present in the general vicinity of Hang Mon in the earliest Miocene.
机译:越南北部杭孟的最早的中新世(阿基坦人,23-21 Ma)杭孟组产生了丰富的陆生腹足纲动物。该组合中的四个物种属于陆足纲腹足类超环纲环线虫。其中三个已分配给东南亚最近的代表属,并被描述为新物种:Cyclophorus hangmonensis Raheem&Schneider sp.。十一月(Cyclophoridae:Cyclophorini),索非斯(Alycaeus sonlaensis)Raheem&Schneider sp.。十一月(Cyclophoridae:Alycaeinae)和Tortulosa naggsi Raheem&Schneider sp。十一月(Pupinidae:Pupinellinae)。这些化石物种代表了其属的最早记录,因此对于校准Cyclophoroidea的分子系统发育具有重要价值。第四个物种仅由保存不佳的片段代表,并保留在环科的开放命名中。虽然现存的Cyclophoroidea在亚洲热带地区具有最大的多样性,但迄今描述的该地区最古老的化石来自晚更新世。杭孟的化石早于这些就存在了两千万年,并且在全球范围内也是当今热带地区记录的最早的环纹石。比中新世更早的Cyclophoroidea仅从欧洲已知。鉴于最近的Cyclophoridae和Pupinidae通常与热带森林有关,因此Hang Mon化石似乎也是热带森林类群,并且在中新世最早出现在Hang Mon的附近地区是合适的栖息地。

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  • 来源
    《Journal of Systematic Palaeontology》 |2018年第15期|1301-1317|共17页
  • 作者单位

    Life Sciences Department, The Natural History Museum, London SW7 5BD, UK;

    CASP, 181A Huntingdon Road, Cambridge CB3 ODH, UK;

    Senckenberg Center for Human Evolution and Paleoecology, University of Tubingen, Institute for Geoscience, Sigwartstrasse 10, 72076 Tubingen, Germany;

    Department of Geosciences, University of Fribourg, Chemin du musee 6, 1700 Fribourg, Switzerland,JURASSICA Museum, Route de Fontenais 21, 2900 Porrentruy, Switzerland;

    Bayerische Staatssammlung fuer Paldontologie und Geologie,Department of Earth and Environmental Sciences, Palaeontology & Geobiology, Ludwig-Maximilians-University Munich, Richard-Wagner-Strasse 10, 80333 Muenchen, Germany;

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  • 原文格式 PDF
  • 正文语种 eng
  • 中图分类
  • 关键词

    Cyclophorus; Alycaeus; Tortidosa; early Miocene; Vietnam; palaeobiogeography;

    机译:Cyclophorus;阿利凯乌斯Tortidosa;中新世早期越南;古生物地理学;

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