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首页> 外文期刊>日本作物學會紀事 >Vegetative Proliferations of Frolal Spikelets in Oryza sativa L. : II. The proliferative structure in the spikelets of the mutant strain induced by X-ray irradiation
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Vegetative Proliferations of Frolal Spikelets in Oryza sativa L. : II. The proliferative structure in the spikelets of the mutant strain induced by X-ray irradiation

机译：玉米苜蓿中的鸡茸小穗的植物增殖。 X射线照射诱导的突变菌株的尖峰中的增殖结构

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The present study was undertaken to determine the fundamental structure of the proliferations common to the "Spikelet-type" having spikelet-like organ(s) in it and the "Leafy-shoot-type" having leafy shoot composed of a number of leafy glumes alternately, and to determine the type of proliferations by the analysis of positional relation between rachilla and the proliferative axis. Results obtained were as follows; 1) The number of empty glumes was increased generally, followed by the increase of the veins in them. 2) Both lower and upper lemma, the latter was corresponding to palea due to its position, had a variety of axillary organs in their axillae, and the frequency of their organ differentiation was increased acropetally. 3) In the half number of the S-type proliferation, the form of the shoot apex was vegetative, showing the transformation into the L-type one. 4) The organs had a 1/2 phyllotaxis in both S-type and L-type proliferations. 5) The L-type proliferation had spikelet-like organs similar to the S-type one at the basal portion of the S-type one. 6) Adventitious roots were recognized at the basal portion of proliferated spikelets and leafy-shoots. As mentioned above, the type of proliferations found in this experiment was recognized to be "diaphysis" which was induced by the apical growth of the shoot apex of rachilla, judging from that the phyllotaxis were the same both in the rachilla and proliferative axis, and the proliferative axis were situated on the extention line of the rachilla and corresponded to the main axis as seen from the upper lemma position. The proliferative structure common to the S-type and the L-type were, therefore, as follows: 1) development of the proliferative axis by apical growth of rachilla differentiating more spikelet-like organs than usual, 2) spikelet-organs were differentiated on the tip of the proliferative axis, 3) their organs were arranged with a 1/2 phyllotaxis from rachilla to the proliferative axis consistently. The proliferation of this experiment, therefore, was grasped as a process of the vegetativation caused by the development of rachilla, and the L-type proliferation was seemed to be a transformed type of the S-type of which the shoot apex continued to differentiate vegetative glumes indefinitely. From the point of the morphogenesis of spikelets, those proliferations were considered to be results of a conversion of growth stage from the differentiation-stage of glumes into that of rachis or spikelet just before the time of differentiation of stamen or pistil. Some discussions were given of the development of the proliferative-body into an independent plantlet, and the signicance of the initiation of adventitious roots.
机译：进行了本研究以确定其具有穗状花序样器官的“穗型型”的增殖的基本结构以及具有由许多叶状葡萄糖组成的叶状芽的“叶状芽型”交替地，通过分析rachilla和增殖轴之间的位置关系来确定增殖的类型。获得的结果如下; 1）空绒布的数量通常增加，其次是静脉增加。 2）均下降和上部引理，后者由于其位置而对应于PALEA，在其腋窝中有各种腋窝器官，并且它们的器官分化的频率增加了致侵扰。 3）在S型增殖的半次数中，枝条的形式是植物派，显示转化为L型。 4）Organs在S型和L型增殖中具有1/2个phyllotaxis。 5）L型增殖具有类似于S型在S型型的S型型型的穗状花套状器官。 6）在增殖小穗和叶芽的基底部分识别不定根。如上所述，在该实验中发现的增殖类型被认为是通过rachilla的枝条的顶端生长诱导的“骨干”，从雷达菌和增殖性轴上判断Phyllotaxis相同，以及增殖性轴位于rAGHILLA的延伸线上，并且与从上部引理位置看到的主轴相对应。因此，S型和L型共同的增殖结构如下：1）通过兔子的顶端生长的增殖轴的发育与往常不同的小穗状器官相差，2）穗器官差异化增殖轴的尖端，3）它们的器官用从兔子始终如一地从rachilla到增殖轴线的1/2个植物。因此，这种实验的增殖被掌握为由rahilla的发育引起的植物的过程，并且L型增殖似乎是转化类型的S型，其中枝条仍继续区分植物无限期地闷闷不乐。从穗穗的形态发生的重点，认为这些增殖是从贫瘠的分化阶段转化生长阶段的结果，进入雄蕊或雌蕊的时间前的rachis或尖峰。一些讨论是给予一个独立的小植物的增殖体的讨论，以及不定根引发的显着性。

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