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首页> 外文期刊>Journal of Systematic Palaeontology >Leaves of Podozamites and Pseudotorellia from the Early Cretaceous of Mongolia: stomatal patterns and implications for relationships
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Leaves of Podozamites and Pseudotorellia from the Early Cretaceous of Mongolia: stomatal patterns and implications for relationships

机译：蒙古早白垩世的假单孢子虫和假杜鹃的叶片：气孔模式及其对关系的影响

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摘要

Strap-shaped, parallel-veined leaves of Podozamites and Pseudotorellia are among the most common fossils in Late Triassic to Early Cretaceous floras from the Northern Hemisphere. Podozamites is considered to be the leaf of a voltzian conifer, but its cuticle is poorly known, whereas Pseudotorellia, with a thick cuticle and haplocheilic stomata, is commonly considered to be the leaf of a ginkgoalean. Here we describe the leaf morphology, cuticle and stomata of Podozamites and Pseudotorellia based on excellently preserved material from the Early Cretaceous of central Mongolia. Podozamites harrisii sp. nov. has transversely oriented, paracytic (probably syndetocheilic) stomata that are regularly arranged in longitudinal files. Pseudotorellia resinosa sp. nov. and Pseudotorellia palustris sp. nov. have scattered, longitudinally oriented stomata in which the two guard cells are sunken and surrounded by 2-5 specialized lateral subsidiary cells and 1-3 unspecialized polar cells. Association evidence and similarities in cuticular structure suggest that Podozamites harrisii was produced by the same plant as the seed cone Krassilovia. The distinctive stomatal pattern of Podozamites harrisii and Krassilovia is also seen in some species of Swedenborgia and Cycadocarpidium, suggesting these plants may all belong to the same natural group. Cycadocarpidium, Krassilovia and Swedenborgia have previously been treated as conifers, but their transversely oriented, paracytic stomata hint instead at a possible relationship with Bennettitales and Gnetales.
机译：带状，平行脉状的Podozamites和Pseudotorellia叶片是北半球晚三叠世至白垩纪早期植物区系中最常见的化石。 Podozamites被认为是沃尔特针叶树的叶子，但是其角质层鲜为人知，而具有厚表皮和单倍气孔的假单胞菌通常被认为是银杏叶的叶子。在这里，我们基于蒙古中部早白垩世保存完好的材料，描述了Podozamites和Pseudotorellia的叶片形态，表皮和气孔。 Podozamites harrisii sp。十一月具有横向排列的，吞噬的（可能是同伴性的）气孔，这些气孔定期排列在纵向文件中。树脂假单胞菌十一月和Pseudotorellia palustris sp.。十一月有散在的，纵向的气孔，其中两个守卫室沉没，并被2-5个专门的侧向辅助细胞和1-3个非专业的极性细胞包围。关联证据和表皮结构的相似性表明，Podozamites harrisii是由与锥锥Krassilovia相同的植物生产的。在一些瑞典紫花and和环卡皮species中也可以看到哈氏Po和克拉斯维亚的气孔模式，这表明这些植物可能都属于同一自然群体。 c果，Krassilovia和Swedishborgia以前被视为针叶树，但它们的横向定向，胞吞气孔暗示了它与Bennettitales和Gnetales的可能关系。

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  • 来源
    《Journal of Systematic Palaeontology》 |2018年第5期|111-137|共27页
  • 作者

    Gongle Shi; Fabiany Herrera; Patrick S. Herendeen; Andrew B. Leslie; Niiden Ichinnorov; Masamichi Takahashi; Peter R. Crane;

  • 作者单位

    "Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing, 210008, China;

    Chicago Botanic Garden, 1000 Lake Cook Road, Glencoe, Illinois 60022, USA;

    Chicago Botanic Garden, 1000 Lake Cook Road, Glencoe, Illinois 60022, USA;

    Department of Ecology and Evolutionary Biology, Brown University, Providence, RI 02912, USA;

    Paleontological Center, Mongolian Academy of Sciences, Ulaanbaatar-51, PO Box 260, Mongolia;

    Department of Environmental Sciences, Faculty of Sciences, Niigata University, 8050, 2-cho, Ikarashi, Nishi-ku, Niigata, 950-2181, Japan;

    School of Forestry and Environmental Studies, Yale University, 195 Prospect Street, New Haven, CT 06511, USA,Oak Spring Garden Foundation, Upperville, VA 20184, USA;

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  • 关键词

    Early Cretaceous; paracytic stoma; Podozamites; voltzian; Pseudotorellia; Ginkgo;

    机译：白垩纪早期;副细胞造口Podozamites;沃尔兹假单胞菌;银杏;

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