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首页> 外文期刊>Journal of Systematic Palaeontology >Septiventeridae, a new and ancestral fossil family of Scarabaeoidea (Insecta: Coleoptera) from the Late Jurassic to Early Cretaceous Yixian Formation
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Septiventeridae, a new and ancestral fossil family of Scarabaeoidea (Insecta: Coleoptera) from the Late Jurassic to Early Cretaceous Yixian Formation

机译：Septiventeridae，从侏罗纪晚期到白垩纪义县早期组的新的祖先化石科Scarabaeoidea（Insecta：Coleoptera）

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摘要

Morphologically, Scarabaeoidea is one of the best-studied groups of beetles. However, the incomplete preservation of presently known fossils is a fundamental problem in the interpretation of extinct species of the superfamily. Wing venation has long been recognized as a valuable character system in taxonomic and phylogenetic analyses. However, to date hind wing features of scarab fossils have not been analysed using geometric morphometrics. A new genus and a new species, Septiventer quadridentatus gen. et sp. nov., is described and illustrated and assigned to a new scarabaeoid family Septiventeridae fam. nov,based on one well-preserved specimen from the Yixian Formation of Liaoning Province, China. The phylogenetic position of Septiventer is inferred based on 68 morphological characters using a cladistic approach. Additionally, based on a geometric morphometric analysis of the hind wing of Septiventer, the structural affinities of 161 scarabs and six outgroup species is analysed, using 261 wing landmarks. Septiventeridae is identified as the sister group of the remaining Scarabaeoidea, with Glaresidae and Trogidae as the next branches. Consequently, it is crucial for an understanding of the early diversification of the superfamily, and for the reconstruction of early evolutionary transformations in the group. Septiventeridae differs strongly from most 'modern' lineages in wing shape. However, the structural affinity of the hind wings of Septiventeridae, Glaresidae and Trogidae are robustly supported by the results from morphometrics. This fits well with the phylogenetic hypothesis based on the general character set and strongly suggests that this wing pattern is closest to the scarabaeoid groundplan. The morphological features suggest good flying abilities of Septiventer, that it might have been active during the daytime, processed soft food, and was less active in digging tunnels than extant, more specialized dung beetles.
机译：从形态上讲，金龟子是研究最多的甲虫之一。但是，目前已知化石的不完全保存是解释超家族灭绝物种的一个基本问题。机翼静脉在分类学和系统发育分析中一直被认为是有价值的特征系统。但是，迄今为止，尚未使用几何形态计量学分析圣甲虫化石的后翅特征。一个新属和一个新物种，Septiventer quadridentatus属。等。 11月进行了描述和说明，并分配给了新的金龟子类Septiventeridae fam。 nov，基于来自中国辽宁省义县组的保存完好的标本。 Seplaventer的系统发育位置是基于68种形态特征，采用分类方法进行推断的。此外，基于对Septiventer后翼的几何形态分析，使用261个翼标，分析了161个圣甲虫和6个外群物种的结构亲和力。 Septiventeridae被确定为剩余的Scarabaeoidea的姊妹组，其中Glaresidae和Trogidae为下一个分支。因此，对于了解超家族的早期多元化以及重建该群体的早期进化转变至关重要。 Septiventeridae与大多数“现代”血统的机翼形状有很大不同。然而，形态计量学的结果有力地支持了七叶科，眩目科和and科的后翅的结构亲和力。这与基于一般特征集的系统发育假说非常吻合，并强烈暗示该机翼模式最接近金龟子纲计划。形态特征表明Septiventer具有良好的飞行能力，它可能在白天活跃，加工软食，并且在挖掘隧道中不如现存的，更专业的虫活跃。

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  • 来源
    《Journal of Systematic Palaeontology》 |2013年第4期|359-374|共16页
  • 作者

    Ming Bai; Rolf G. Beutel; Chung-Kun Shih; Dong Ren; Xing-Ke Yang;

  • 作者单位

    Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing 100101, PR China;

    Institut fuer Spezielle Zoologie und Evolutionsbiologie, FSU Jena,Germany;

    College of Life Sciences, Capital Normal University, Beijing 100048, PR China;

    College of Life Sciences, Capital Normal University, Beijing 100048, PR China;

    Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Box 92, No. 1, Beichen West Road, Chaoyang District, Beijing 100101, PR China;

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  • 关键词

    Septiventeridae; Septiventer; phylogeny; geometric morphometries; new family; fossil;

    机译：科Septiventer;系统发育几何形态新家庭化石;

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