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Tuzoia: Morphology and lifestyle of a large bivalved arthropod of the Cambrian Seas

机译:Tuzoia:寒武纪大型双壳节肢动物节肢动物的形态和生活方式

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The morphology of Titzoia is reinterpreted in the light of abundant new specimens from the Middle Cambrian Burgess Shale (British Columbia, Canada) and Kaili (Guizhou, China) Lagerstatten. Tuzoia was a very large (up to 180 mm long) bivalved arthropod with a nonmineralized domelike carapace strengthened by prominent pointed features and often flanked by a lateral ridge bearing a spiny frill. The reticulate pattern of Tuzoia is comparable with that of present-day crustaceans (e.g., myodocope ostracods) and is interpreted as a structural compromise between exoskeletal lightness and high resistance to mechanical stress. Tuzoia had a pair of large. stalked, spherical, possibly compound eyes facing forward. Flagella-like antennae protruded through the anterior notch. No other appendages are known except possible filamentous setae underlying the carapace. Tuzoia typically occurs as laterally (l) or dorsoventrally (dvc) compacted carapaces or single valves. Each type (lc or dvc) emphasizes particular aspects of the morphology (e.g., spiny lateral ridge. ventral margin) that were often interpreted as specific differences by previous authors. A revision of Tuzoia validates only 7 of the 23 named species. Tuzoia is placed tentatively within a group of large bivalved arthropods along with Isoxys and the possible ancestors of Thylacocephala (Lower Cambrian-Upper Cretaceous). In the Middle Cambrian, Titzoia occurs across Laurentia, South and North China, and the Perigondwanan area (Bohemia) within a relatively narrow subtropical belt, indicating a high dispersal capability and possible latitudinal control on its distribution. Functional morphology, taphonomy, and the distributional pattern indicate that Tuzoia was a free-swimming arthropod.
机译:通过从中寒武纪伯吉斯页岩(加拿大不列颠哥伦比亚省)和凯里市(中国贵州省)拉格施塔滕的大量新标本重新解释了Titzoia的形态。 Tuzoia是一个非常大(长至180毫米长)的双壳节肢动物节肢动物,具有非矿化的圆顶状甲壳,通过突出的尖锐特征得到加强,并且通常两侧为带有多刺褶边的侧脊。 Tuzoia的网状图案可与当今甲壳类动物(例如,Myodocope ostracods)相媲美,并且被解释为骨骼外的轻度和对机械应力的高抵抗力之间的结构折衷。 Tuzoia有一对大的。茎秆呈球形,可能是复眼,朝前。鞭毛状触角穿过前切口突出。除了在甲壳下面的可能的丝状刚毛以外,没有其他附件。裂口虫通常以侧向(l)或背腹(dvc)压实的甲壳或单瓣出现。每种类型(lc或dvc)都强调形态的特定方面(例如,棘突侧脊,腹缘),以前的作者通常将其解释为特定差异。 Tuzoia的修订版仅验证了23个命名物种中的7个。 Tuzoia被暂时放置在一组大型双壳节肢动物节肢动物中,与Isoxys和Thylacocephala(下寒武统-上白垩统)的可能祖先一起。在中寒武纪,Titzoia遍布劳伦西亚,华南和华北以及亚热带相对较窄带内的Perigondwanan地区(波希米亚),表明其扩散能力高,并且可能对其分布进行纬度控制。功能形态,拓扑学和分布模式表明,Tuzoia是自由游泳的节肢动物。

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