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RELATIONSHIPS AMONG FOLIAR PHENOLOGY, RADIAL GROWTH RATE, AND XYLEM DENSITY IN A YOUNG DOUGLAS-FIR PLANTATION

机译:道格拉斯松人工林中叶片物候,径向生长速率和木糖密度的关系

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摘要

We related intra-annual patterns in radial growth rate and xylem density to foliar phenology and second growth flushes in a young Douglas-fir plantation in western Washington. Three foliar maturity classes were defined: (1) shoots and needles elongating; (2) elongation complete, needles maturing; and (3) needles mature. Diameter growth rate had two peaks, one about the time of budbreak and one when foliage was in maturity Class 2. There was a limit to the maximum periodic density of xylem formed at agiven rate of diameter growth; as growth rate increased, maximum periodic density decreased. Although xylem density profiles varied widely among individual trees, xylem density differed significantly among foliar maturity classes, increasing 16% from Class 1 to 2 and 60% from Class 2 to 3. Diameter growth rate of second-flushing trees was significantly greater in July compared with trees with no second flush, but we detected no relationships between second-flushing and xylem density patterns or false rings. Although the young trees in this study did not show distinct earlywood-latewood transitions, fully mature foliage (Class 3) was associated with formation of xylem exhibiting characteristics of latewood: greater density, reduced diameter growth rate, reduced tracheid radial diameter, and less interannual growth variation.
机译:我们将华盛顿西部一个年轻的道格拉斯冷杉人工林中径向生长速率和木质部密度的年内模式与叶片物候和第二次生长潮红联系起来。定义了三个叶类的成熟度类别:(1)枝条和针伸长; (2)伸长完成,针头成熟; (3)针头成熟。直径的生长有两个峰值,一个是在萌芽时出现的,另一个是在叶子达到成熟等级2时出现的。在给定的直径增长速率下,形成的木质部的最大周期性密度是有限的。随着增长率的增加,最大周期性密度降低。尽管各个树木的木质部密度分布差异很大,但叶成熟度类别之间木质部密度差异显着,从1级到2级增加了16%,从2级到3级增加了60%。没有第二次冲洗的树木,但是我们没有检测到第二次冲洗与木质部密度模式或假环之间的关系。尽管本研究中的幼树没有表现出明显的早木-晚木过渡,但完全成熟的树叶(3类)与形成具有晚木特征的木质部有关:密度更高,直径增长速度降低,气管径向直径降低且年际更少增长差异。

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